Starscream's History
Starscream is easily as accomplished, powerful and deserving of leadership as Megatron. At least, that's what he thinks. Power-hungry and driven by jealousy, Starscream operates through three ideals: lying, manipulation and deception. And when all else fails, a bit of sabotage never goes astray. Wise bots are careful about turning their backs on Starscream, just in case it's the day he finally decides to shove in the knife.
Though not as powerful as Megatron (despite what he thinks of himself), Starscream is by no means a wimp. His weapons can demolish large buildings, and he can fly around so fast that he can break free of Ratchet's magnetic powers. Unfortunately, all these abilities are offset by an enormous ego and the tendency to make long, overbearing speeches when he should just get on with it and fight. More than once, his leanings towards bombast have caused him to miss a golden opportunity. Sadly, he doesn't seem to be learning from these mistakes.
During the Great War, Starscream destroyed Theta Supreme while he was being towed away for repairs.
Starscream was aboard the Nemesis when it came across an Autobot repair ship that appeared to be emitting the energy signature of the fabled AllSpark. Starscream's doubt of the AllSpark's very existence and his dismissal of Megatron's millennia-long search did not sit well with Lugnut, and an argument broke out on the bridge until Megatron himself arrived and all fell silent. Starscream was forced to eat his words when Blackarachnia confirmed that the energy coming off the ship was indeed that of the AllSpark, but quickly volunteered to secure it for his leader. Seeing through his lieutenant's greed, Megatron put a dent in Starscream's wing and rose to obtain it for himself. As he was leaving, however, Starscream arrived at the airlock to wish Megatron luck and patted him on the back, planting a shatter bomb on him in the process. As Megatron attacked the Autobot ship, the device overloaded his circuits and caused an explosion that led Starscream to believe that Megatron was no more. In the aftermath of the blast, Starscream got so caught up in his own greatness that he didn't notice that his comrades had left him on the ship, hurtling towards an exploding space bridge.
Surviving the explosion, Starscream spent the next fifty solar cycles searching for the AllSpark on his own, using the damaged ship as a base (and probably getting a few screws loose in his central processor in the meantime). Detecting the faint signal of the AllSpark on Earth, Starscream tapped into television broadcasts from the planet and discovered that the Autobot ship had landed there.
While heading for the planet, Starscream was approached by Blackarachnia, who was also seeking the AllSpark for her own purposes and proposed an alliance. Starscream was unsurprisingly unwilling to share power, and he booted Blackarachnia out into space just before the ship warped away. Starscream was so occupied by the task, however, that he failed to notice Earth's moon appearing in the ship's path
One crash-landing later, Starscream abandoned the ruined hulk of the ship and headed down to Earth, immediately drawing the attention of military jets. After a brief dogfight, Starscream scanned one of the jets as an alternate mode in order to continuing searching incognito. The search did not go on long, as Starscream found the Autobots watching a Crimson Angels air show, and he quickly joined in the proceedings. Upon revealing himself to the Autobots, he quickly thrashed each of them individually, mocking their belief that they had defeated "the great Megatron" while boldly admitting to planting the bomb. Though his arrogance initially trumped his physical power, a sustained blast allowed Starscream to claim the AllSpark. Optimus Prime managed to recover and grabbed Starscream in midair. The two's struggle for the AllSpark opened its container, unleashing a burst of its energy that sent Starscream hurtling off.